Supervised and unsupervised deep learning-based approaches for studying DNA replication spatiotemporal dynamics.

Journal: Communications biology
PMID:

Abstract

In eukaryotic cells, DNA replication is organised both spatially and temporally, as evidenced by the stage-specific spatial distribution of replication foci in the nucleus. Despite the genetic association of aberrant DNA replication with numerous human diseases, the labour-intensive methods employed to study DNA replication have hindered large-scale analyses of its roles in pathological processes. In this study, we employ two distinct methodologies. We first apply supervised machine learning, successfully classifying S-phase patterns in wild-type mouse embryonic stem cells (mESCs), while additionally identifying altered replication dynamics in Rif1-deficient mESCs. Given the constraints imposed by a classification-based approach, we then develop an unsupervised method for large-scale detection of aberrant S-phase cells. Such a method, which does not aim to classify patterns based on pre-defined categories but rather detects differences autonomously, closely recapitulates expected differences across genotypes. We therefore extend our approach to a well-characterised cellular model of inducible deregulated origin firing, involving cyclin E overexpression. Through parallel EdU- and PCNA-based analyses, we demonstrate the potential applicability of our method to patient samples, offering a means to identify the contribution of deregulated DNA replication to a plethora of pathogenic processes.

Authors

  • Julian Ng-Kee-Kwong
    Institute of Cell Biology, School of Biological Sciences, University of Edinburgh, Roger Land Building, Alexander Crum Brown Road, Edinburgh, EH9 3FF, UK.
  • Ben Philps
    School of Informatics, University of Edinburgh, Edinburgh, EH8 9AB, UK.
  • Fiona N C Smith
    School of Informatics, University of Edinburgh, Edinburgh, EH8 9AB, UK.
  • Aleksandra Sobieska
    School of Informatics, University of Edinburgh, Edinburgh, EH8 9AB, UK.
  • Naiming Chen
    Institute of Cell Biology, School of Biological Sciences, University of Edinburgh, Roger Land Building, Alexander Crum Brown Road, Edinburgh, EH9 3FF, UK.
  • Constance Alabert
    Division of Molecular, Cell & Developmental Biology, School of Life Sciences, University of Dundee, Dundee, DD15EH, UK.
  • Hakan Bilen
    School of Informatics, University of Edinburgh, Edinburgh, EH8 9AB, UK.
  • Sara C B Buonomo
    Institute of Cell Biology, School of Biological Sciences, University of Edinburgh, Roger Land Building, Alexander Crum Brown Road, Edinburgh, EH9 3FF, UK. sara.buonomo@ed.ac.uk.