Structured networks support sparse traveling waves in rodent somatosensory cortex.

Journal: Proceedings of the National Academy of Sciences of the United States of America
Published Date:

Abstract

Neurons responding to different whiskers are spatially intermixed in the superficial layer 2/3 (L2/3) of the rodent barrel cortex, where a single whisker deflection activates a sparse, distributed neuronal population that spans multiple cortical columns. How the superficial layer of the rodent barrel cortex is organized to support such distributed sensory representations is not clear. In a computer model, we tested the hypothesis that sensory representations in L2/3 of the rodent barrel cortex are formed by activity propagation horizontally within L2/3 from a site of initial activation. The model explained the observed properties of L2/3 neurons, including the low average response probability in the majority of responding L2/3 neurons, and the existence of a small subset of reliably responding L2/3 neurons. Sparsely propagating traveling waves similar to those observed in L2/3 of the rodent barrel cortex occurred in the model only when a subnetwork of strongly connected neurons was immersed in a much larger network of weakly connected neurons.

Authors

  • Samat Moldakarimov
    Howard Hughes Medical Institute, The Salk Institute for Biological Studies, La Jolla, CA 92037.
  • Maxim Bazhenov
    Department of Cell Biology and Neuroscience, University of California at Riverside, Riverside, California 92521 bazhenov@salk.edu.
  • Daniel E Feldman
    Department of Molecular and Cell Biology, University of California, Berkeley, CA 94720.
  • Terrence J Sejnowski
    Howard Hughes Medical Institute, Salk Institute for Biological Studies, La Jolla, United States.